A

Abel, D. C., Grubbs, R. D. (Profesores). (2018). MSCI 473/473L: Shark Biology (Curso). Coastal Carolina University.

Abel, D. C., Grubbs, R. D. (Profesores). (2021). MSCI 473/473L: Shark Biology (Curso). Coastal Carolina University.

Abel, D. C., Grubbs, R. D. (2020). Shark biology and conservation. Johns Hopkins University Press. 

Adams, L., Lyons, K., Monday, J., Larkin, E., Wyffels, J. (2023). Costs of parthenogenesis on growth and longevity in ex situ zebra sharks Stegostoma tigrinum. Endangered Species Research, 50, 81–91. https://doi.org/10.3354/esr01224

Agencia de Protección Ambiental de Estados Unidos (2023, noviembre 16). Información básica sobre el sargazo. US EPA. https://espanol.epa.gov/espanol/informacion-basica-sobre-el-sargazo#:~:text=Podemos%20encontrar%20dos%20especies%20de,tallos%20cortos%20y%20hojas%20anchas.

Aines, A. C., Carlson, J. K., Boustany, A., Mathers, A., Kohler, N. E. (2017). Feeding Habits of the Tiger Shark, Galeocerdo cuvier, in the Northwest Atlantic Ocean and Gulf of Mexico. Environmental Biology of Fishes, 101(3). https://doi.org/10.1007/s10641-017-0706-y

Alicea Segarra, D. del C., Bejarano Rodríguez, I., Soler Figueroa, B. M., Casillas Maldonado, J. I., Martínez, H. M. (2016). Los arrecifes de coral: Guía educativa para maestros. Programa de Colegio Sea Grant de la Universidad de Puerto Rico.

Alicea Segarra, D. del C., Casillas Maldonado, J. I., Bejarano Rodríguez, I. (2014). Las praderas de hierbas marinas: Guía educativa para maestros. Programa de Colegio Sea Grant de la Universidad de Puerto Rico.

All in One Fishing Charters (2019). What Sharks are in the SC Waters Series? https://www.allinonecharters.com/what-sharks-are-in-the-sc-waters-series-learn-about-blacknose-sharks/

Amigos de los Dinosaurios y la Paleontología. (2022, septiembre 18). Era del Carbonífero. Mundo Prehistórico. https://www.mundoprehistorico.com/carbonifero/

Anderson, R. I. (2025). ESOblog: Probing Polaris ESO Fellow Richard I. Anderson sheds light on the famous North Star. European Southern Observatory. https://www.eso.org/public/blog/probing-polaris/#:~:text=It’s%20actually%20a%20multiple%20system,more%20than%20500%20times%20fainter).

Animals Network Team (2018). Lionfish. Animals.NET. https://animals.net/lionfish/

Aquatic Pathobiology Laboratory (s.f.). Fish Guts: A Guide to Fish Anatomy, Health, and Necropsy. University of Florida. https://aquaticpath.phhp.ufl.edu/fg2/anat/jobs.html#:~:text=Gill%20rakers%20are%20an%20anatomical,is%20pictured%20on%20the%20right

Artisanal Fisheries. (2015). American Bluefin Tuna Association. https://www.theabta.com/artisanal

Artisanal Fisheries: Meaning, Characteristics & Dangers. (2022). Ocean Scientists. https://www.oceanscientists.org/artisanal-fisheries/

Asunsolo-Rivera, A. Lester, E., Langlois, T., Vaughan, B. I., McCormick, M. I., Simpson, S. D., Meekan, M. G. (2023). Behaviour of mesopredatory coral reef fishes in response to threats from sharks and humans. Scientific Reports, 13(1). https://doi.org/10.1038/s41598-023-33415-5

Atlantic Highly Migratory Species (HMS) Fisheries; Fishery Management Plan (FMP), Plan Amendment, and Consolidation of Regulations of 1999, 16 U.S.C. 971 et seq.; 16 U.S.C. 1801 et seq. (1999). https://www.ecfr.gov/current/title-50/chapter-VI/part-635

B

Báez Rodríguez, N., Harms-Tuohy, C., Schizas, N. V. (2022). Using DNA Barcoding to Identify Seafood Fraud in Puerto Rico. Caribbean Journal of Science, 52(2). https://doi.org/10.18475/cjos.v52i2.a14

Baker Jr., M. S., Sciance, M. B., Halls, J. N. (2016). Potential for a Simple GPS-based Binary Logit Model to Predict Fishing Effort in a Vertical Hook-and-Line Reef Fish Fishery. Marine and Coastal Fisheries, 8, 118-131. https://doi.org/10.1080/19425120.2015.1135221

Barber, M., Mitchell, W., von Hirsch, T., Vyas, T. (2021). A Drop in the Ocean: Closing the Gap in Ocean Climate Finance. Deloitte. https://www2.deloitte.com/content/dam/Deloitte/uk/Documents/risk/ocean-financing.pdf

Barker, A. M., Frazier, B., Gelsleichter, J., Grubbs, R. D., Hollenbeck, C. M., Portnoy, D. (2019). High Rates of Genetic Polyandry in the Blacknose Shark, Carcharhinus acronotus. Copeia, 107(3), 502-508. https://doi.org/10.1643/CG-19-180

BD Editors (2017). Taxonomy. Biology Dictionary. https://biologydictionary.net/taxonomy/

Begum, T. (2023, febrero 21). What is mass extinction and are we facing a sixth one? Natural History Museum. https://www.nhm.ac.uk/discover/what-is-mass-extinction-and-are-we-facing-a-sixth-one.html

Bennet, M. V. L. (1971). Electrolocation in Fish. Annals of the New York Academy of Sciences. 

Bergen, M. (2016). 5 Things You Didn’t Know Sharks Do for You. Conservation International. https://www.conservation.org/blog/5-things-you-didnt-know-sharks-do-for-you#:~:text=Sharks%20help%20keep%20the%20carbon,move%20carbon%20through%20the%20ocean

Berkovitz, B., Shellis, P. (2017). The Teeth of Non-Mammalian Vertebrates. Academic Press.

Bernardini, I., Garibaldi, F., Canesi, L., Fossi, M. C., Baini, M. (2018). First data on plastic ingestion by blue sharks (Prionace glauca) from the Ligurian Sea (North-Western Mediterranean Sea). Marine Pollution Bulletin, 135, 303–310. doi:10.1016/j.marpolbul.2018.07.022

Bird of prey. (2013). DBpedia. https://dbpedia.org/page/Bird_of_prey

Bittel, J. (2017). Alligators Attack and Eat Sharks, Study Confirms. National Geographic. https://www.nationalgeographic.com/animals/article/american-alligator-sharks-predators-stingrays

Black, R. (2012, diciembre 5). Scientists Discover Oldest Known Dinosaur. Smithsonian Magazine. https://www.smithsonianmag.com/science-nature/scientists-discover-oldest-known-dinosaur-152807497/

Bluntnose sixgill shark. (2025). Washington Department of Fish & Wildlife. https://wdfw.wa.gov/species-habitats/species/hexanchus-griseus#:~:text=Bluntnose%20sixgill%20sharks%20can%20grow,due%20to%20large%20litter%20size.

Boca de río. (2018). Glosbe.  https://es.glosbe.com/es/es/boca%20de%20r%C3%ADo

Bond, M. E., Valentin-Albanese, J., Babcock, E. A., Hussey, N. E., Heithaus, M. R., Chapman, D. D. (2018). The trophic ecology of Caribbean reef sharks (Carcharhinus perezi) relative to other large teleost predators on an isolated coral atoll. Marine Biology, 165(4). https://doi.org/10.1007/s00227-018-3322-2

Bonfil, R., Clarke, S., Nakano, H. (2008). The Biology and Ecology of the Oceanic Whitetip Shark, Carcharhinus longimanus. En M. D. Camhi, E. K. Pikitch, E. A. Babcock (Eds.), Sharks of the Open Ocean, 128–139. Blackwell Publishing Ltd. doi:10.1002/9781444302516.ch11

Brown, A. M. (2004). The impacts of dredging and habitat destruction on the lemon shark, Negaprion brevirostris, population of Bimini, Bahamas. doi: 10.13140/RG.2.2.16149.32488.

Bunkley-Williams, L., Williams, E. (2004). New locality, depth, and size records and species character modifications of some Caribbean deep-reef/ shallow slope fishes and a new host and locality record for the chimaera ces- todarian. Caribbean Journal of Science, 40, 88-119.

C

Cailliet, G. M., Musick, J. A., Simpfendorfer, C. A., Stevens, J. D. (2005). Ecology and Life History Characteristics of Chondrichthyan Fish. In Sharks, Rays and Chimeras:The Status of Chondrychthyan Fish, Status Survey. IUCN / Shark Specialist Group: IUCN, Gland, Switzerland and Cambridge, UK. 

Camhi, M. D., Fowler, S., Musick, J., Bräutigam, A., Fordham, S. (1998). Sharks and their Relatives: Ecology and Conservation. The IUCN Species Survival Commission.

Camhi, M. D., Pikitch, E. K., Babcock, E. A. (Eds.) (2008). Sharks of the Open Ocean: Biology, Fisheries and Conservation. Blackwell Publishing Ltd.

Campana, S. E., Marks, L., Joyce, W., Kohler, N. E. (2006). Effects of Recreational and Commercial Fishing on Blue Sharks (Prionace glauca) in Atlantic Canada, with Inferences on the North Atlantic Population. Canadian Journal of Fisheries and Aquatic Sciences, 63, 670 – 682. 

Cardeñosa, D.,Shea, S. K., Zhang, H., Fischer, G. A., Simpfendorfer, C. A., Chapman, D. A. (2022). Two thirds of species in a global shark fin trade hub are threatened with extinction: Conservation potential of international trade regulations for coastal sharks. Conservation Letters, 15, e12910. https://doi.org/10.1111/conl.12910 

Caribbean Fishery Management Council (2025, noviembre 30). ¡En Veda la Cabrilla! https://www.facebook.com/caribbeanfishery/posts/𝐂𝐨𝐦𝐢𝐞𝐧𝐳𝐚-𝐞𝐥-𝟏-𝐝𝐞-𝐝𝐢𝐜𝐢𝐞𝐦𝐛𝐫𝐞-𝐥𝐚-𝐯𝐞𝐝𝐚-𝐝𝐞𝐥-𝐦𝐞𝐫𝐨-𝐜𝐚𝐛𝐫𝐢𝐥𝐥𝐚-𝐞𝐧-𝐚𝐠𝐮𝐚𝐬-𝐥𝐨𝐜𝐚𝐥𝐞𝐬-𝐝𝐞-𝐏𝐮𝐞𝐫𝐭𝐨-/1284216950412360/

Caribbean Fishery Management Council, National Marine Fisheries Service (2023). Amendment 3 to the Puerto Rico, St. Croix, and St. Thomas and St. John Fishery Management Plans: Management Measures for Dolphin and Wahoo. https://issuu.com/caribbeanfmc/docs/draft_amendment_3_dolphin_and_wahoo_management_mea 

Carrasco de la Cruz, P. M. (2021). The Knowledge Status of Coastal and Marine Ecosystem Services – Challenges, Limitations and Lessons Learned From the Application of the Ecosystem Services Approach in Management. Frontiers in Marine Science, 13(8). https://doi.org/10.3389/fmars.2021.684770

Carrier, J. C., Simpfendorfer, C. A., Heithaus, M. R., Yopak, K. E. (Eds.). (2022). Biology of Sharks and their Relatives. CRC Press.

Carruba, L., Pabón, A., O’Callaghan, J., Lugo, M., López, M., Blackinship, R., Ortiz, L., Pemberton, R., Tobias, W., Jiménez, N., Rosario, A., Alcover-Serrano, I., Esteves-Amador, R. [Eds.]. (2016). Regulated and Protected Species in U.S. Caribbean Waters: Invertebrates, Fish, Sea Turtles, Marine Mammals. NOAA Coral Reef Conservation Program: Coral Reef Information System. http://www.coris.noaa.gov/activities/protectedSpecies/ 

Castro, J. I. (2013). Historical Knowledge of Sharks: Ancient Science, Earliest American Encounters, and American Science, Fisheries, and Utilization. Marine Fisheries Review, 75 (4), 1-26.

Castro, J. I. (2009). Observations on the reproductive cycles of some viviparous North American sharks. aqua, International Journal of Ichthyology, 15(4), 205-222.

Castro, J. (2011). The Sharks of North America. Oxford University Press.

Castro, J. I., Wourms, J. P. (1993). Reproduction, placentation, and embryonic development of the Atlantic sharpnose shark, Rhizoprionodon terraenovae. Journal of Morphology, 218(3), 257–280. doi:10.1002/jmor.1052180304

Catéter urinario: Cuidado en casa. (2019). The Hospital for Sick Children. https://www.aboutkidshealth.ca/article?contentid=1246&language=spanish

Catlin, J., Hughes, M., Jones, T., Jones, R., Campbell, R. (2013). Valuing Individual Animals Through Tourism: Science or Speculation? Biological Conservation, 157, 93–98. doi:10.1016/j.biocon.2012.07.022

Chang, C.-H., Kao, Y.-T., Huang, T.-T., & Wang, Y.-C. (2021). Product Authentication Using Two Mitochondrial Markers Reveals Inconsistent Labeling and Substitution of Canned Tuna Products in the Taiwanese Market. Foods, 10(11), 2655. https://doi.org/10.3390/foods10112655

Chapman, D. D., Firchau, B., Shivji, M. S. (2008). Parthenogenesis in a large-bodied requiem shark, the blacktip Carcharhinus limbatus. Journal of Fish Biology, 73 (6), 1473–1477. https://doi.org/10.1111/j.1095-8649.2008.02018.x

Cherney, K. (2018). Understanding Mercury Poisoning. Healthline. https://www.healthline.com/health/mercury-poisoning#symptoms

Cisneros-Montemayor, A. M., Barnes, M. L., Dalal Al-Abdulrazzak, Navarro‐Holm, E. E., Sumaila, U. R. (2013). Global economic value of shark ecotourism: implications for conservation. Oryx, 47(3), 381–388. https://doi.org/10.1017/s0030605312001718

CITES. (2023). Appendices. https://cites.org/eng/app/appendices.php

Claeson. K. M. (2011) The synarcual cartilage of batoids with emphasis on the synarcual of Rajidae. Journal of Morphology, 272(12), 1444-63. doi: 10.1002/jmor.10996. 

Claeson, K. M., Dean, M.N. (2011). The Skeleton: Cartilaginous Fish Skeletal Anatomy. En A. P. Farrel (Ed.), Encyclopedia of Fish Physiology, 419-427. Academic Press.

Cleveland Clinic (2022, mayo 23). Collagen: What It Is, Types, Function & Benefits. https://my.clevelandclinic.org/health/articles/23089-collagen

Collin, S. P. (2021). Scene through the Eyes of an Apex Predator: A Comparative Analysis of the Shark Visual System. Clinical and Experimental Optometry, 101(5), 624-640. https://doi.org/10.1111/cxo.12823

Comité de Estándares y Peticiones de la UICN (2019). Directrices de uso de las Categorías y Criterios de la Lista Roja de la UICN. Versión 14. https://www.iucnredlist.org/es/resources/redlistguidelines

Common Cuttlefish. (2023). Monterey Bay Aquarium. https://www.montereybayaquarium.org/animals/animals-a-to-z/common-cuttlefish

Commonwealth of Puerto Rico Department of Natural and Environmental Resources Supplementary Flier Puerto Rico Fishing Regulations. (2004). https://www.caribbeanfmc.com/meetings/CFMC%20MEETINGS/127%20regular%20meeting-MARCH%2026-27-2008/127%20presentaciones/PR%20Fishing%20Regs%20-%20good%20one.pdf

Compagno L. J. V. (1984). Vol. 4: Sharks of the World: An Annotated and Illustrated Catalogue of Shark Species Known to Date Part 2. Food and Agriculture Organization. https://www.fao.org/3/ad123e/ad123e35.pdf

Conrath, C. L., Musick, J. A. (2002). Reproductive Biology of the Smooth Dogfish, Mustelus canis, in the Northwest Atlantic Ocean. Environmental Biology of Fishes, 64(4), 367–377. doi:10.1023/a:1016117415855

Consumo de “cecina de tiburón” un peligro para la salud. (2013). OSPESCA. https://www.sica.int/busqueda/Noticias.aspx?IDItem=76612&IDCat=2&IdEnt=47

Cook, S. (Ed.). (1987). Sharks: An Inquiry into Biology, Behavior, Fisheries and Use. Oregon State University Extension Service. 

Correa de Carvalho, A. P., Rangel, B., Ferreira de Amorim, A. (2020). Insights into the reproductive biology of sharpnose sevengill shark (Heptranchias perlo) in the western South Atlantic. En E. da Silva (Ed.), Tópicos Multidisciplinares em Ciências Biológicas, 2, 45-60. Atena Editora.

Cortés, E. (2000). Life History Patterns and Correlations in Sharks. Reviews in Fisheries Science, 8(4), 299-344.

Cortés, E., Brooks, E. N., Shertzer, K. W. (2015). Risk Assessment of Cartilaginous Fish Populations. ICES Journal of Marine Science, 72 (3), 1057 – 1068. 

Cortés, E., Domingo, A., Miller, P., Forselledo, R., Mas, F., Arocha, F., Campana, S., Coelho, R., Da Silva, C., Hazin, F. H. V., Holtzhausen, H., Keene, K., Lucena, F., Ramirez, K., Santos, M. N., Semba-Murakami, Y., Yokawa, K. (2015). Expanded Ecological Risk Assessment of Pelagic Sharks Caught in Atlantic Pelagic Longline Fisheries. ICAAT, 71 (6), 2637 – 2688. 

Cortés, E., Arocha, F., Beerkricher, L, Carvalho, F., Domingo, A., Heupel, M., Holtzhausen, H., Santos, M. N., Ribera, M., Simpfendorfer, C. (2010). Ecological Risk Assessment of Pelagic Sharks Caught in Atlantic Pelagic Longline Fisheries. Aquatic Living Resources, 23, 25 – 34.  

Costa, M. E., Borges, T. C., Capapé, C. (2013). Cases of abnormal hermaphroditism in velvet belly Etmopterus spinax (Chondrichthyes: Etmopteridae) from the southern coast of Portugal. Cahiers de Biologie Marine, 54(3), 309-317.

CSIRO (2017). Shark Litters Have Multiple Dads According to DNA Tests. CSIROscope. https://blog.csiro.au/sharkpaternity/#:~:text=%E2%80%9CWe%20now%20know%20that%20multiple,two%20and%20eight%20different%20fathers

D

Daly-Engel, T., Baremore, I. E., Grubbs, R. D., Gulak, S. J. B., Graham, R. T., Enzenauer, M. P. (2018). Resurrection of the sixgill shark Hexanchus vitulus Springer & Waller, 1969 (Hexanchiformes, Hexanchidae), with comments on its distribution in the northwest Atlantic Ocean. Marine Biodiversity. https://doi.org/10.1007/s12526-018-0849-x

Danovaro, R., Aguzzi, J., Fanelli, E., Billett, D., Gjerde, K., Jamieson, A., Ramirez-Llodra, E., Smith, C. R., Snelgrove, P. V. R., Thomsen, L., Van Dover, C. L. (2017). An Ecosystem-Based Deep-Ocean Strategy. Science, 355(6324), 452-454. doi: 10.1126/science.aah7178

Dennis, G. D. (2000). Annotated checklist of shallow-water marine fishes from the Puerto Rico Plateau including Puerto Rico, Culebra, Vieques, St. Thomas, St. John, Tortola, Virgin Gorda, and Anegada; U.S. Geological Survey, Florida Caribbean Science Center. https://doi.org/10.5281/zenodo.802287

Departamento de Recursos Naturales y Ambientales (2023). Laboratorio de Investigaciones Pesqueras. https://www.drna.pr.gov/oficinas/laboratorio-pesquero/

Departamento de Recursos Naturales y Ambientales de Puerto Rico. (2004). Reglamento de Pesca de Puerto Rico. http://app.estado.gobierno.pr/ReglamentosOnLine/Reglamentos/6768.pdf

De Vos, L. (2021). What Eats sharks? Save Our Seas Foundation. https://saveourseas.com/worldofsharks/what-eats-sharks

Díaz, I. (2023, octubre). Guía completa para calcular la abundancia relativa en ecología. Ecología Digital. https://ecologiadigital.bio/como-se-calcula-la-abundancia-relativa-en-ecologia-y-que-importancia-tiene/

Dillman, C. (2018). CUMV Fish Collection. Version 28.16. Cornell University Museum of Vertebrates. Occurrence dataset. https://doi.org/10.15468/jornbc recuperado de GBIF.org https://www.gbif.org/occurrence/3698456658

Dudgeon, C. L., Corrigan, S., Yang, L., Allen, G. R., Erdmann, M. V., Fahmi, Hagi Yulia Sugeha, White, W. T., Naylor, G. J. P. (2020). Walking, swimming or hitching a ride? Phylogenetics and biogeography of the walking shark genus Hemiscyllium. Marine and Freshwater Research, 71(9), 1107–1107. https://doi.org/10.1071/mf19163

Duffy, L. M., Griffiths, S. P. (2019). Assessing attribute redundancy in the application of productivity-susceptibility analysis to data-limited fisheries. Aquatic Living Resources, 32, 20–20. https://doi.org/10.1051/alr/2019018

Dulvy, N. K., Fowler, S. L., Musick, J. A., Cavanagh, R. D., Kyne, P. K., Harrison, L. R., Carlson, J. K., Davidson, L. N.K., Fordham, S. V., Francis, M. P., Pollock, C. M., Simpfendorfer, C. A., Burgess, G. H., Carpenter, K. E., Compagno, L. J.V., Ebert, D. A., Gibson, C., Heupel, M. R., Livingstone, S. R., Sanciangco, J. C., Stevens, J. D., Valenti, S., White, W. T. (2014). Extinction Risk and Conservation of the World’s Sharks and Rays. eLife, 3. 

Dulvy N. K., Pacoureau, N., Rigby, C. L., Pollom, R. A., Jabado, R. W., Ebert, D. A., Finucci, B., Pollock, C. M., Cheok, J., Derrick, D. H., Herman, K. B., Sherman, C. S., VanderWright, W. J., Lawson, J. M., Walls, R. H. L., Carlson, J. K., Charvet, P., Bineesh, K. K., Fernando, D., Ralph, G. M., Matsushiba, J. H., Hilton-Taylor, C., Fordham, S. V., Simpfendorfer, C. A. (2021). Overfishing drives over one-third of all sharks and rays toward a global extinction crisis. Current Biology, 31, 1-15.

E

Ebert, D. A., Dando, M., Fowler, S. (2021). Sharks of the World: A Complete Guide. Princeton University Press.

Ebert, D. A., Stehmann, M. F. W. (2013). Sharks, Batoids and Chimaeras of the North Atlantic. Food and Agriculture Organization of the United Nations.

Ectotérmico (2023). Etimologías de Chile. https://etimologias.dechile.net/?ectote.rmico

El Colegio de México (2026). Desovar. Diccionario del Español de México. https://dem.colmex.mx/

Eliassen, Meredith, MSLIS (2023). Western Culture. EBSCO. https://www.ebsco.com/research-starters/history/western-culture#western-culture

Enax, J., Prymak, O., Raabe, D., Epple, M. (2012). Structure, composition, and mechanical properties of shark teeth. Journal of Structural Biology, 178(3), 290–299. doi:10.1016/j.jsb.2012.03.012

Endangered Species Act of 1973, Pub. L. No. 93-205, 87 Stat. 884 (codified as amended at 16 U.S.C. §§ 1531–1544 (2018)).

Endosos y Permisos (2018). Secretaría Auxiliar de Permisos, Endosos y Servicios Especializados. Departamento de Recursos Naturales y Ambientales, Gobierno de Puerto Rico. https://www.drna.pr.gov/oficinas/sap/#:~:text=Descripci%C3%B3n,unidades%20que%20conforman%20el%20Departamento.

Engelbrecht, T. M., Kock, A. A., O’Riain, M. J. (2019). Running Scared: When Predators Become Prey. Ecosphere, 10(1), e02531. https://doi.org/10.1002/ecs2.2531

Erdman, D. S. (1983). Nombres Vulgares de los Peces en Puerto Rico. CODREMAR.

Escamas ctenoideas. (2020). Biodiversidad Virtual. https://www.biodiversidadvirtual.org/taxofoto/glosario/1564

Especies de fauna silvestre. (2018, febrero). Argentina.gob.ar. https://www.argentina.gob.ar/ambiente/biodiversidad/fauna-silvestre#:~:text=La%20fauna%20silvestre%2C%20tambi%C3%A9n%20llamada,en%20forma%20permanente%2C%20circunstancial%20o

F

Fadool, B. A., Bostick, K. G., Brewster, L. R., Hansell, A. C., Carlson, J. K., Smukall, M. J. (2024). Age and growth estimates for the nurse shark (Ginglymostoma cirratum) over 17 years in Bimini, The Bahamas. Frontiers in Marine Science, 11. https://doi.org/10.3389/fmars.2024.1265150

Fake Declared Shark Meat. (2017). VisionDive. https://www.visiondive.com/deceptive-names-for-shark-meat/

Farlex, INC (2023). The Free Dictionary. https://www.thefreedictionary.com/

Fedler, A. J. (2017). The Economic Impact of Shark Diving in Florida. Oceana. https://usa.oceana.org/wp-content/uploads/sites/4/economic_impact_of_shark_diving_in_florida_report.pdf

Feichtinger, I., Lukeneder, A., Topa, D., Kriwet, J., Libowitzky, E., Westall, F. (2020). Fossil microbial shark tooth decay documents in situ metabolism of enameloid proteins as nutrition source in deep water environments. Scientific Reports, 10, 20979. https://doi.org/10.1038/s41598-020-77964-5

Feldheim, K. A., Chapman, D. D., Sweet, D., Fitzpatrick, S., Prodöhl, P. A., Shivji, M. S., Snowden, B. (2010). Shark Virgin Birth Produces Multiple, Viable Offspring. Journal of Heredity, 101(3), 374–377. https://doi.org/10.1093/jhered/esp129

Feldheim, K. A., Gruber, S. H., Dibattista, J. D., Babcock, E. A., Kessel, S. T., Hendry, A. P., Pikitch, E. K., Ashley, M. V., Chapman, D. D. (2014). Two Decades of Genetic Profiling Yields First Evidence of Natal Philopatry And Long-Term Fidelity to Parturition Sites in Sharks. Molecular Ecology, 23(1), 110-117. https://doi.org/10.1111/mec.12583

Ferretti, F., Worm, B., Britten, G. L., Heithaus, M. R., Lotze, H. K. (2010). Patterns and Ecosystem Consequences of Shark Declines in the Ocean. Ecology Letters, 13, 1055-1071. 

Finucci, B., Pacoureau, N., Rigby, C. L., Matsushiba, J. H., Faure-Beaulieu, N., Sherman, C. S., VanderWright, W. J., Jabado, R. W., Charvet, P., Mejía-Falla, P. A., Navia, A. F., Derrick, D. H., Kyne, P. M., Pollom, R. A., Rachel, Herman, K. B., Bineesh Kinattumkara, Cotton, C. F., Cuevas, J.-M., Daley, R. K. (2024). Fishing for oil and meat drives irreversible defaunation of deepwater sharks and rays. Science, 383(6687), 1135–1141. https://doi.org/10.1126/science.ade9121

Fischer, A. (2022). El tiburón ballena podría desaparecer a fuerza de chocar contra embarcaciones industriales. National Geographic en Español. https://www.ngenespanol.com/animales/el-tiburon-ballena-podria-desaparecer-por-embarcaciones-industriales/

Florida Museum of Natural History (2023). Discover Fishes. University of Florida. https://www.floridamuseum.ufl.edu/

Florida Museum of Natural History (2023). International Shark Attack File. University of Florida. https://www.floridamuseum.ufl.edu/

Florida Museum of Natural History (2023). Learn About Sharks. University of Florida. https://www.floridamuseum.ufl.edu/

Fontes, J., McGinty, N., Machete, M., Afonso, P. (2020). Whale Shark-Tuna Associations, Insights from a Small Pole-and-Line Fishery from the Mid-North Atlantic. Fisheries Research, 229, 105598. https://doi.org/10.1016/j.fishres.2020.105598 

Food and Agriculture Organization of the United Nations. (2018). The State of World Fisheries and Aquaculture 2018: Meeting the Sustainable Development Goals.

Fourqurean, J. W., Duarte, C. M., Kennedy, H., Marbà, N., Holmer, M., Mateo, M. A., Apostolaki, E. T., Kendrick, G. A., Krause-Jensen, D., McGlathery, K. J., Serrano, O. (2012). Seagrass Ecosystems as a Globally Significant Carbon Stock. Nature Geoscience, 5, 505–509. https://doi.org/10.1038/ngeo1477

Fowler, S. L., Cavanagh, R. D., Camhi, M., Burgess, G. H., Cailliet, G. M., Fordham, S. V., Simpfendorfer, C. A., Musick, J. A. (Eds.). (2005). Sharks, Rays and Chimeras: The Status of Chondrichthyan Fishes. IUCN/SSC Shark Specialist Group. https://portals.iucn.org/library/efiles/documents/2005-029.pdf

Fradera vda. de Hernández, V. (2011). Peces de Puerto Rico y de las Islas Vírgenes EUA. Programa de Colegio Sea Grant de la Universidad de Puerto Rico.

Franqui-Rivera, G., Rodriguez-Ferrer, G., Schizas, N. (2025) Collaborative effort between fishers and scientists reveals first-time evidence of the smalltooth sandtiger shark (Odontaspis ferox) in Puerto Rico – CORRIGENDUM. Journal of the Marine Biological Association of the United Kingdom, 105, e99, 1. https://doi.org/10.1017/ S0025315425100271

G 

Gall, S.C., Thompson, R.C. (2015). The impact of debris on marine life. Marine Pollution Bulletin, 92(1-2), 170–179. doi:10.1016/j.marpolbul.2014.12.041

Gallagher, Austin J., Hammerschlag, N. (2011). Global shark currency: the distribution, frequency, and economic value of shark ecotourism. Current Issues in Tourism, 14(8), 797–812. doi:10.1080/13683500.2011.585227

Gallagher, A. J., Hammerschlag, N., Danylchuk, A. J., Cooke, S. J. (2017). Shark Recreational Fisheries: Status, Challenges, and Research Needs. Ambio, 46, 385 – 398. 

Gallagher, A. J., Kyne, P. M., Hammerschlag, N. (2012). Ecological Risk Assessment and Its Application to Elasmobranch Conservation and Management. Journal of Fish Biology, 80(5), 1727-1748.

García, V. B., Lucifora, L. O., Myers, R. A. (2008). The Importance of Habitat and Life History to Extinction Risk in Sharks, Skates, Rays and Chimaeras. Proceedings of of the Royal Society B, 275, 83-89. doi: 10.1098/rspb.2007.1295

Geib, C. (2017). For Sharks, Even Catch and Release Can Kill. Hakai Magazine. https://hakaimagazine.com/news/sharks-even-catch-and-release-can-kill/#:~:text=But%20what%20happens%20after%E2%80%94once,still%20die%20from%20the%20stress. 

Gemballa, S., Konstantinidis, P., Donley, J. M., Sepulveda, C., Shadwick, R. E. (2006). Evolution of High-Performance Swimming in Sharks: Transformations of the Musculotendinous System from Subcarangiform to Thunniform Swimmers. Journal of Morphology, 267 (4), 477-493. doi:10.1002/jmor.10412

Genetic material (2021). Bioloogy Online. https://www.biologyonline.com/dictionary/genetic-material

Germain, J. (2022). Steven Spielberg Regrets How “Jaws” Impacted Real-World Sharks. Smithsonian Magazine. https://www.smithsonianmag.com/smart-news/steven-spielberg-regrets-how-jaws-impacted-real-world-sharks-180981335/

Gibbens, S. (2017). Extremely Rare Megamouth Shark Filmed. National Geographic. https://www.nationalgeographic.com/animals/article/rare-megamouth-shark-sighting-video-spd

Gilbert, P. W. (1962). The Behavior of Sharks. Scientific American, 207(1), 60-69. https://www.jstor.org/stable/10.2307/24936606

Gilmore, R. G. (1993). Reproductive Biology of Lamnoid Sharks. Environmental Biology of Fishes, 38(1-3), 95–114. doi:10.1007/bf00842907

Gobierno de Puerto Rico. (2020). Ley de Pesquerías de Puerto Rico. https://bvirtualogp.pr.gov/ogp/Bvirtual/leyesreferencia/PDF/Recursos%20Naturales/278-1998/278-1998.pdf

Goh, R., Yoshida, E., Schaible, E., Behrens, R., Monnier, C. A., Killingsworth, B., Kong, K. W., Hiew, S. H., Miserez, A., Hoon, S., Waite, J. H. (2022). Nanolattice-Forming Hybrid Collagens in Protective Shark Egg Cases. Biomacromolecules, 23(7), 2878–2890. https://doi.org/10.1021/acs.biomac.2c00341

González, M. (2020). Key Reef Herbivores of Puerto Rico. Departamento de Recursos Naturales y Ambientales de Puerto Rico. https://www.drna.pr.gov/wp-content/uploads/2020/11/Final-Key-Reef-Herbivores-COMPRESSED_compressed.pdf

González-Pestana, A., Acuña-Perales, N., Córdova, F., Coasaca, J., Alfaro, E., Alfaro-Shigueto, J., Mangel, J. (2019). Feeding Habits Of Thresher Sharks Alopias sp. in Northern Peru: Predators of Humboldt Squid (Dosidicus gigas). Journal of the Marine Biological Association of the United Kingdom, 99 (3), 695-702. doi:10.1017/S0025315418000504

Grace, M. (2001). Field Guide to Requiem Sharks (Elasmobranchiomorphi: Carcharhinidae) of the Western North Atlantic, NOAA Technical Report NMFS 153: A Scientific Paper of the Fishery Bulletin. U.S. Department of Commerce.

Grana Raffucci, F. A. (2007). Nomenclatura de los organismos acuáticos y marinos de Puerto Rico e Islas Vírgenes. Departamento de Recursos Naturales y Ambientales de Puerto Rico. 

Gravinese, P. M., Perry, S. A., Angelo Jason Spadaro, Boyd, A. E., Enochs, I. C. (2022). Caribbean king crab larvae and juveniles show tolerance to ocean acidification and ocean warming. Marine Biology, 169(5). https://doi.org/10.1007/s00227-022-04053-8

Greenaway, S. F., Sullivan, K. D., Harper Umfress, S., Beittel, A. B., Wagner, K. D. (2021). Revised depth of the Challenger Deep from submersible transects; including a general method for precise, pressure-derived depths in the ocean. Deep-Sea Research I, 178, 10364. https://doi.org/10.1016/j.dsr.2021.103644

Gress, E., Andradi-Brown, D. A., Woodall, L., Schofield, P. J., Stanley, K., Rogers, A. D. (2017). Lionfish (Pterois spp.) Invade the Upper-Bathyal Zone in the Western Atlantic. PeerJ Life & Environment, 5, e3683. doi: 10.7717/peerj.3683

Grupo de Innovación sobre la Docencia en Diversidad Biológica (2022). Mecanismos de termorregulación. BIOINNOVA. https://www.innovabiologia.com/biodiversidad/diversidad-animal/mecanismos-de-termorregulacion/#:~:text=El%20proceso%20que%20controla%20la,(con%20temperatura%20corporal%20variable).

Guerrero Pérez, C. R., García, M. A., Lilyestrom, C., Rodríguez, G., Rodríguez, Y. (2014). Puerto Rico Marine Recreational Fisheries Statistics Program: Final Report 2009 – 2013. Department of Natural and Environmental Resources: Fish and Wildlife Bureau: Marine Resources Division.

Gulf of Maine Research Institute (2019). Marine Resource Education Program. https://mrep.gmri.org/

Guttridge, T. L., Gruber, S. H., Gledhill, K. S., Croft, D. P., Sims, D. W., Krause, J. (2009). Social preferences of juvenile lemon sharks, Negaprion brevirostris. Animal Behaviour, 78(2), 543–548. doi:10.1016/j.anbehav.2009.06.009

H

Haas, A. R., Fedler, T., Brooks, E. J. (2017). The contemporary economic value of elasmobranchs in The Bahamas: Reaping the rewards of 25 years of stewardship and conservation. Biological Conservation, 207, 55–63. https://doi.org/10.1016/j.biocon.2017.01.007

Hall, B. K. (2015). Bones and Cartilage. Academic Press.

Hall, D. (2023, May 11). Ocean Through Time. Smithsonian Ocean; Smithsonian National Museum of Natural History. https://ocean.si.edu/through-time/ocean-through-time

Hamlett, W. C. (Ed.) (1999). Sharks, Skates, and Rays: The Biology of Elasmobranch. Johns Hopkins University Press.

Handwerk, B. (2021, febrero 2). An Evolutionary Timeline of Homo Sapiens. Smithsonian Magazine. https://www.smithsonianmag.com/science-nature/essential-timeline-understanding-evolution-homo-sapiens-180976807/#:~:text=The%20long%20evolutionary%20journey%20that,in%20their%20tools%20and%20technologies.

Haque, A. B., Cavanagh, R. D., Seddon, N. (2021). Evaluating artisanal fishing of globally threatened sharks and rays in the Bay of Bengal, Bangladesh. PLoS ONE, 16(9), e0256146. https://doi.org/10.1371/journal.pone.0256146

Harms-Tuohy, C. A., Schizas, N. V., Appeldoorn, R. S. (2016). Use of DNA Metabarcoding for Stomach Content Analysis in the Invasive Lionfish Pterois volitans in Puerto Rico. Marine Ecology Progress Series, 558, 181-191. doi: 10.3354/meps11738

Heavy Metal Poisoning (Heavy Metal Toxicity): Symptoms, Causes & Treatment. (2022). Cleveland Clinic. https://my.clevelandclinic.org/health/diseases/23424-heavy-metal-poisoning-toxicity

Heim, V., Dhellemmes, F., Smukall, M. J., Gruber, S. H., Guttridge, T. L. (2021) Effects of Food Provisioning on the Daily Ration and Dive Site Use of Great Hammerhead Sharks, Sphyrna mokarran. Frontiers in Marine Science, 8(628469). doi: 10.3389/fmars.2021.628469

Heithaus, M. R., Frid, A., Wirsing, A. J., Worm, B. (2007). Predicting Ecological Consequences of Marine Top Predator Declines. Trends in Ecology and Evolution, 23(4), 202-209. 

Heupel, M. R., Carlson, J. K., Simpfendorfer, C. A. (2007). Shark Nursery Areas: Concepts, Definition, Characterization and Assumptions. Marine Ecology Progress Series, 337, 287-297.

Histotrophy. (2022). En Wikipedia. https://en.wikipedia.org/wiki/Histotrophy

Horn, C., Karnauskas, M., Doerr, J., Miller, M., Neuman, M., Hill, R., & Mccarthy, K. (2022). Endangered Species Act Status Review Report: Queen Conch (Aliger gigas). https://doi.org/10.25923/4ykr-1m56

Horstmyer, L., Ballard, K., Daly-Engel, T., Stein, K., Hoffmayer, E., Hammerschlag, N., Frazier, B. S., Anderson, B., Campbell, B., Sulikowski, J. (2025). First insights into the fine-scale vertical movements of a Carolina hammerhead, Sphyrna gilberti, and a hybrid between Carolina and scalloped hammerhead. Frontiers in Marine Science, 12. https://doi.org/10.3389/fmars.2025.1688202

Howard, B. C. (2019). These Sharks Glow Underwater—Thanks to Tiny ‘Lightsabers’. National Geographic. https://www.nationalgeographic.com/animals/article/biofluorescent-catsharks-glow-underwater-with-tiny-lightsabers-metabolites

How Jaws Influenced Shark Perception. (2023). Shark Stewards. https://sharkstewards.org/how-jaws-influenced-shark-perception/#:~:text=The%201975%20film%20Jaws%20had,popular%20culture%20to%20this%20day.

Hung, R. (2015). Fish Biology and Fish Scales. Rs’ Science. https://rsscience.com/fish-biology-and-fish-scales-under-the-microscope/

Hunt, C. L., Andradi-Brown, D. A., Hudson, C. J., Bennett-Williams, J., Noades, F., Curtis-Quick, J., Lewis, O. T., Exton, D. A. (2020). Shelter Use Interactions of Invasive Lionfish with Commercially and Ecologically Important Native Invertebrates on Caribbean Coral Reefs. PLoS One, 15(8), e0236200. https://doi.org/10.1371/journal.pone.0236200

Hussey, N. E., DiBattista, J. D., Moore, J. W., Ward, E. J., Fisk, A. T., Kessel, S., Guttridge, T. L., Feldheim, K. A., Franks, B. R., Gruber S. H., Weideli, O. C., Chapman, D. D. (2017). Risky business for a juvenile marine predator? Testing the influence of foraging strategies on size and growth rate under natural conditions. Proceedings of the Royal Society B: Biological Sciences. https://doi.org/10.1098/rspb.2017.0166

I

Inner Space Center (2013). E/V Nautilus Video Bite: 6 Gill Shark Encounter – Puerto Rico 2013 [Video]. YouTube. https://www.youtube.com/watch?v=00SOMpc8diw.

J

Jabado, R. W., Morata, A. Z. A., Bennett, R. H., Finucci, B., Ellis, J. R., Fowler, S., Grant, M. I, Barbosa Martins, A. P., Sinclair, S. L. (2024). The global status of sharks, rays, and chimaeras. IUCN: IUCN Library System. https://portals.iucn.org/library/node/52102

Jamieson, A. J., Stewart, H. A., Nargeolet, P. H. (2020). Exploration of the Puerto Rico Trench in the Mid-twentieth Century: Today’s Significance and Relevance. Endeavour, 44(1–2). https://doi.org/10.1016/j.endeavour.2020.100719

Jennings, D. E., Gruber, S. H., Franks, B. R., Kessel, S. T., Robertson, A. L. (2008). Effects of large-scale anthropogenic development on juvenile lemon shark (Negaprion brevirostris) populations of Bimini, Bahamas. Environmental Biology of Fishes, 83, 369–377. https://doi.org/10.1007/s10641-008-9357-3

Jerome, R. (Escritor), Kennedy, K. (Director de edición) (2020). JAWS: A Legacy of Fear. Life. 

Jolly, D. (2011). Priced Off the Menu? Palau’s Sharks Are Worth $1.9 Million Each, a Study Says. The New York Times. https://www.nytimes.com/2011/05/02/science/earth/02shark.html

K

Kalmijn, A. J. (1988). Detection of Weak Electric Fields. Sensory Biology of Aquatic Animals, 151–186. doi:10.1007/978-1-4612-3714-3_6

Keartes, S. (2022). Whale Sharks Team Up to Hunt with Other Predators in Rare Video. National Geographic. https://www.nationalgeographic.com/animals/article/whale-sharks-can-hunt-with-other-sharks-feed-on-bait-balls

Keartes, S. (2020). Do Sharks Bite People? Save Our Seas Foundation. https://saveourseas.com/worldofsharks/why-do-sharks-bite-people

Kemp, N. E. (1977). Banding Pattern and Fibrillogenesis of Ceratotrichia in Shark Fins, Journal of Morphology, 154, 187-204.

Kennedy, E., Xu, D., Mankad, R., Heaps, L., Midgley, A., Holmes, L., de Liedekerke, V. (2021). Navigating Ocean Risk. WWF, Metabolic. 

Kimura, S., Kubota, M. (1969). Tyrosin Derivatives in a Structural Protein, Elastoidin. The Journal of Biochemistry, 65(1).

Kiraly, S. J., Moore, J. A., Jasinski, P. H. (2003). Deepwater and Other Sharks of the U.S. Atlantic Ocean Exclusive Economic Zone. Marine Fisheries Review, 65(4).

Knipp, J. (2019). What is Neutral Buoyancy and How to Achieve Perfect Neutral Buoyancy While Diving? Scuba Guru. https://scubaguru.org/nj-scuba-articles/what-is-neutral-buoyancy-and-how-to-achieve-perfect-neutral-buoyancy-while-diving/

Kohler, N. E., Turner, P. A. (2019). Distributions and Movements of Atlantic Shark Species: A 52-Year Retrospective Atlas of Mark and Recapture Data. Marine Fisheries Review, 8(12).

Konstantinou, H., Cozzi, J. R. (1998). Galeus springeri, a New Species of Sawtail Catshark from the Caribbean Sea (Chondrichthyes, Scyliorhinidae). Copeia, 1998(1), 151-158. doi:10.2307/1447711

Kyne, P. M., Davies, C.-L., Floriaan Devloo‐Delva, Johnson, G., Yolarnie Amepou, Grant, M. I., Green, A., Gunasekera, R. M., Harry, A. V., Lemon, T., Lindsay, R., Maloney, T., Marthick, J. R., Pillans, R. D., Saunders, T., Shields, A., Shields, M., Feutry, P. (2021). Molecular analysis of newly-discovered geographic range of the threatened river shark Glyphis glyphis reveals distinct populations. Charles Darwin University; CSIRO. https://researchers.cdu.edu.au/en/publications/molecular-analysis-of-newly-discovered-geographic-range-of-the-th/

L

La mayor amenaza para los tiburones es el consumo de su carne en Europa y América Latina. (2018). Notimerica. https://www.notimerica.com/sociedad/noticia-mayor-amenaza-tiburones-consumo-carne-europa-america-latina-20180215141038.html

Larson, S., Christiansen, J., Griffing, D., Ashe, J., Lowry, D., Andrews, K. (2011). Relatedness and Polyandry of Sixgill Sharks, Hexanchus griseus, in an Urban Estuary. Conservation Genetics, 12, 679–690. https://doi.org/10.1007/s10592-010-0174-9

Learn, J. R. (2018, mayo 22). Historical Art Paints a Picture of Past Shark Abundance. Hakai Magazine. https://hakaimagazine.com/features/historical-art-paints-picture-past-shark-abundance/#:~:text=In%20492%20BCE%2C%20the%20first,the%20work%20of%20visual%20artists.

Leigh, S. C., Papastamatiou, Y. P., German, D. P. (2018). Seagrass digestion by a notorious “carnivore.” Proceedings of the Royal Society B: Biological Sciences, 285(1886), 20181583–20181583. https://doi.org/10.1098/rspb.2018.1583

Leigh S. C., Summers A. P.,Hoffmann S. L., German D. P. (2021). Shark Spiral Intestines May Operate as Tesla Valves. Proceedings of the Royal Society B: Biological Sciences, 288, 20211359. https://doi.org/10.1098/rspb.2021.13

Lester, E., Meekan, M. G., Barnes, P., Raudino, H. C., Rob, D., Waples, K., Speed, C. W. (2020). Multi-Year Patterns in Scarring, Survival and Residency of Whale Sharks in Ningaloo Marine Park, Western Australia. Marine Ecology Progress Series, 634, 115–125. https://doi.org/10.3354/meps13173

Ley Núm. 93 del año 2025: Ley para el Manejo de Especies Introducidas. (2025). LexJuris Puerto Rico. https://www.lexjuris.com/lexlex/Leyes2025/lexl2025093.htm

Ley Núm. 295 del año 2004 – Ley para enmendar los Arts. 2 y 4 de la Ley Núm. 241 de 1999: Nueva Ley de Vida Silvestre de Puerto Rico. (2025). LexJuris Puerto Rico. https://www.lexjuris.com/lexlex/Leyes2004/lexl2004295.htm

Linnaeus NG (2023). Sharks of the World. Naturalis Biodiversity Center. https://linnaeus.naturalis.nl

Long, J. A., Trinajstic, K. (2024). Placoderms. Current Biology, 34 (2), R43–R47. https://doi.org/10.1016/j.cub.2023.11.038

Longnecker, M. P., Rogan, W. J., Lucier, G. W. (1997). The Human Health Effects of DDT (Dichlorodiphenyltrichloroethane) And PCBs (Polychlorinated Biphenyls) and an Overview of Organochlorines in Public Health. Annual Review of Public Health, 18(1), 211–244. https://doi.org/10.1146/annurev.publhealth.18.1.211

Lowry, B. (2014). Hakarl: Traditional Iceland Fermented Shark. The Foodie Blog; Culinary Schools. https://www.culinaryschools.org/blog/hakarl/

Lozano-Grande, M. A., Gorinstein, S., Espitia-Rangel, E., Dávila-Ortiz, G., Martínez-Ayala, A. L. (2018). Plant Sources, Extraction Methods, and Uses of Squalene. International Journal of Agronomy, 2018(1829160), 1–13. https://doi.org/10.1155/2018/1829160

LUMITOS AG (2023). Vejiga natatoria. QUIMICA.ES. https://www.quimica.es/enciclopedia/Vejiga_natatoria.html#:~:text=El%20proceso%20de%20secreci%C3%B3n%20gaseosa,difunden%20en%20la%20vejiga%20natatoria.

Lyon, B. J., Dwyer, R. G., Pillans, R. D., Campbell, H. A., Franklin, C. E. (2017). Distribution, seasonal movements and habitat utilisation of an endangered shark, Glyphis glyphis, from northern Australia. Marine Ecology Progress Series, 573, 203-213. https://doi.org/10.3354/meps12200

M

Mallefet, J., Stevens, D. W., Duchatelet, L. (2021). Bioluminescence of the Largest Luminous Vertebrate, the Kitefin Shark, Dalatias licha: First Insights and Comparative Aspects. Frontiers in Marine Science, 8. https://doi.org/10.3389/fmars.2021.633582

Manard, B. T., Hintz, C. J., C. Derrick Quarles, Burns, W. K., N. Alex Zirakparvar, Dunlap, D. R., Beiswenger, T., Cruz-Uribe, A. M., Petrus, J. A., Hexel, C. R. (2022). Determination of fluorine distribution in shark teeth by laser-induced breakdown spectroscopy. Metallomics, 14(7). https://doi.org/10.1093/mtomcs/mfac050

Mariani, G., Cheung, W. W. L., Lyet, A., Sala, E., Mayorga, J., Velez, L., Gaines, S. D., Dejean, T., Troussellier, M., Mouillot, D. (2020). Let More Big Fish Sink: Fisheries Prevent Blue Carbon Sequestration—Half in Unprofitable Areas. Science Advances, 6(44). doi:10.1126/sciadv.abb4848

Marine Conservation Wildlife (2022). Basking Shark Guide: How Big They Are, What They Eat, and Why They’re Endangered. Discover Wildlife. https://www.discoverwildlife.com/animal-facts/fish/facts-about-basking-sharks/

Marine Stewardship Council (2026). What is a fishery. Marine Stewardship Council: Australia & New Zealand. https://www.msc.org/en-au/what-we-are-doing/our-collective-impact/what-is-a-fishery#:~:text=A%20basic%20definition%20of%20a,purpose%20of%20the%20activities.

Martin. R. A. (2003). A Golden Agen of Sharks. ReefQuest Center for Shark Research. http://www.elasmo-research.org/education/evolution/golden_age.htm

R. A. (2003). Deep Sea: Beyond the Twillight Zone: Cookiecutter Shark. ReefQuest Center for Shark Research. http://www.elasmo-research.org/education/ecology/deepsea-cookiecutter.htm

Márquez, M. C. (Profesora). (2022). The Truth About Sharks (Curso). Atlas Obscura.

Martin, R. A. (2005). Conservation of Freshwater and Euryhaline Elasmobranchs: A Review. Journal of the Marine Biological Association of the UK, 85(5), 1049-1073. 

Matos Caraballo, D. (2012). Puerto Rico/NMFS Cooperative Fisheries Statistics Program April 2007 – September 2012. Department of Natural and Environmental Resources.

Mattos, S. M. G., Broadhurst, M., Hazin, F. H. V., Jonnes, D. M. (2001). Reproductive biology of the Caribbean sharpnose shark, Rhizoprionodon porosus, from northern Brazil. Marine and Freshwater Research, 52(5), 745-752.

Mazhar, M. (2021, junio 28). Scientists find world’s oldest evidence of a shark attack — and it isn’t pretty. CBC Radio. https://www.cbc.ca/radio/asithappens/scientists-find-world-s-oldest-evidence-of-a-shark-attack-and-it-isn-t-pretty-1.6083041#:~:text=According%20to%20a%20study%20from%20the%20University,to%20those%20caused%20by%20a%20tiger%20shark.

McCauley, D. J., McLean, K. A., Bauer, J., Young, H. S., Micheli, F. (2012). Evaluating the Performance of Methods for Estimating the Abundance of Rapidly Declining Coastal Shark Populations. Ecological Applications, 22 (2), 385 – 392. 

McCoy, M. K. (2023). Indonesia protects ‘walking sharks.’ Are other sharks next? Conservation International. https://www.conservation.org/blog/indonesia-protects-walking-sharks-are-other-sharks-next

Meekan, M. G., Virtue, P., Marcus, L., Clements, K. D., Nichols, P. D., & Revill, A. T. (2022). The world’s largest omnivore is a fish. Ecology, 103(12). https://doi.org/10.1002/ecy.3818

Mercury Cycle. (2023). Woods Hole Oceanographic Institution. https://www.whoi.edu/multimedia/mercury-cycle/

Merriam-Webster, INC (2023). Merriam-Webster Dictionary. https://www.merriam-webster.com/

Messner, G. F. (1990). The Shark-Calling Ceremony in Paruai, New Ireland, Papua New Guinea. The World of Music, 32(1), 49-83. https://www.jstor.org/stable/43561243

Meyer, C. G., Dale, J. J., Papastamatiou, Y. P., Whitney, N. M., Holland, K. N. (2009). Seasonal Cycles and Long-Term Trends in Abundance and Species Composition of Sharks Associated with Cage Diving Ecotourism Activities in Hawaii. Environmental Conservation, 36(2), 104 – 111.

Mignucci-Giannoni, A. A., Cintrón-Nieves, D., Franqui-Rivera, G., Espinoza, R., Orcera-Iglesias, J. M., Rivera-Ilarraza, P. J., Rivera-Pérez, C. I., Rodríguez-Ferrer, G. (2020). First Confirmed Record of the Longfin Mako Shark (Isurus paucus) for Puerto Rico. Caribbean Naturalist, 71, 1-9.

MI News Network (2022). 10 Deepest Parts of the Ocean. Marine Insight. https://www.marineinsight.com/know-more/10-deepest-parts-of-the-ocean/

Ministerio de la Producción (2020). Produce: ¿Qué es una veda y para qué sirve? Gob.pe. https://www.gob.pe/institucion/produce/noticias/312623-produce-que-es-una-veda-y-para-que-sirve

Miloslavich, P., Díaz, J. M., Klein, E., Alvarado, J. J., Díaz, C., Gobin, J., Escobar-Briones, E., Cruz-Motta, J. J., Weil. E., Cortés, J., Bastidas, A. C., Robertson, R., Zapata, F., Martín, A., Castillo, J., Kazandjian, A., Ortiz, M. (2010). Marine Biodiversity in the Caribbean: Regional Estimates and Distribution Patterns. PLoS One, 5 (8).

Mojetta, A. R., Travaglini, A., Scacco, U., Bottaro, M. (2018). Where sharks met humans: The Mediterranean Sea, history and myth of an ancient interaction between two dominant predators. Regional Studies in Marine Science, 21, 30–38. https://doi.org/10.1016/j.rsma.2017.10.001

Molles Jr., M. C. (2008). Ecology Concepts & Applications. McGraw-Hill Higher Education. 

Morris, J. A., Akins, J. L. (2009). Feeding Ecology of Invasive Lionfish (Pterois volitans) in The Bahamian Archipelago. Environmental Biology of Fishes, 86(3), 389–398. doi:10.1007/s10641-009-9538-8

Murray, R. W. (1974). The Ampullae of Lorenzini. In Electroreceptors and Other Specialized Receptors in Lower Vertebrates.

Musick, J. A., Burgess, G., Cailliet, G., Camhi, M., Fordham, S. (2000). Management of Sharks and Their Relatives (Elasmobranchii). AFS Policy Statement. 

Musick, J. A. (1999). Ecology and Conservation of Long-Lived Marine Animals. American Fisheries Society Symposium, 23, 1-10. 

N

Nakaya, K., William Toby White,  Hsuan Ching, H. (2020). Discovery of a new mode of oviparous reproduction in sharks and its evolutionary implications. Scientific Reports, 10, 12280. doi:10.1038/s41598-020-68923-1

Nakamura, I., Meyer, C. G., Sato K. (2015). Unexpected Positive Buoyancy in Deep Sea Sharks, Hexanchus griseus, and a Echinorhinus cookei. PLoS ONE, 10(6), e0127667. doi:10.1371/journal.pone.0127667

NASA Space Place (2019, junio 27). How Does GPS Work? NASA Science. https://spaceplace.nasa.gov/gps/en/#:~:text=Satellites%20act%20like%20the%20stars,to%20within%20a%20few%20inches!

National Aquarium (2023). Nurse Shark. https://aqua.org/explore/animals/nurse-shark#:~:text=Females%20produce%20a%20litter%20of,bull%20and%20great%20hammerhead%20sharks.

National Cancer Institute (2023). Small Intestine. En Dictionary of Cancer Terms. National Institutes of Health. https://www.cancer.gov/publications/dictionaries/cancer-terms/def/small-intestine

National Cancer Institute (2026). Ultraviolet radiation. En Dictionary of Cancer Terms. National Institutes of Health. https://www.cancer.gov/publications/dictionaries/cancer-terms/def/ultraviolet-radiation

National Geographic Education. (2023, octubre 19). Marine Food Pyramid. National Geographic. https://education.nationalgeographic.org/resource/marine-food-pyramid-1/

National Human Genome Research Institute. (2026). Bacteria. National Institutes of Health. https://www.genome.gov/genetics-glossary/Bacteria

National Oceanic and Atmospheric Administration. (2025, septiembre 30). Marine mammals. https://www.noaa.gov/education/resource-collections/marine-life/marine-mammals

National Ocean Service (2023). Coral Reefs Support Jobs, Tourism, and Fisheries. Florida Keys National Marine Sanctuary. https://floridakeys.noaa.gov/corals/economy.html

National Ocean Service (2023). Invasive Lionfish. Flower Garden Banks National Marine Sanctuary. https://flowergarden.noaa.gov/education/invasivelionfish.html

National Ocean Service (2015). What are Microplastics? NOAA. https://oceanservice.noaa.gov/facts/microplastics.html

National Ocean Service (2023). What are PCBs? NOAA. https://oceanservice.noaa.gov/facts/pcbs.html

National Ocean Service (2023). What is a coral reef made of? NOAA. https://oceanservice.noaa.gov/facts/coralmadeof.html

National Ocean Service (2015). What Is a Lionfish? NOAA. https://oceanservice.noaa.gov/facts/lionfish-facts.html

National Ocean Service (2023). What Is an Invasive Species? NOAA. https://oceanservice.noaa.gov/facts/invasive.html

Naylor, G., Florida Program for Shark Research, Florida Museum of Natural History (2023). Chondrichthyan Tree of Life. University of Florida.  https://sharksrays.org/

Neilson, H. R., Blinn, H. (2020). The Curious Case of the North Star: the continuing tension between evolution models and measurements of Polaris. ArXiv.org. https://arxiv.org/abs/2003.02326

Nelson, J. S., Grande, T. C., Wilson, M. V. H. (2016). Fishes of the World. John Wiley & Sons.

New Hampshire PBS (2023). Chordata: Vertebrates, Tunicates, Lancelets. Wildlife Journal Junior. https://nhpbs.org/wild/chordata.asp#:~:text=All%20chordates%20have%20a%20post,lancelets%2C%20tunicats%2C%20and%20vertebrates.

Nicoletti, A. (2022). High levels of toxic mercury in some species of shark meat, fins pose dangers to human health. FIU News. https://news.fiu.edu/2022/mercury

Nilsson, H. C., Valentinsson, D. (2023). Selective Fishing for Sustainable Use of Marine Ecosystems. SLU; Swedish University of Agricultural Sciences. https://www.slu.se/en/departments/aquatic-resources1/selective-fishing/

NOAA Fisheries. (2024). Understanding Fisheries Management in the United States. NOAA. https://www.fisheries.noaa.gov/insight/understanding-fisheries-management-united-states#:~:text=The%20U.S.%20fisheries%20management%20process%20includes:%20,Implementing%20gear%20restrictions%20%20Protecting%20sensitive%20habitats

NOAA Fisheries (2023). Endangered Species Act. NOAA. https://www.fisheries.noaa.gov/national/endangered-species-conservation/endangered-species-act

NOAA Fisheries (2023). Permiso Comercial Caribeño de Botes Pequeños para HMS: Requisitos de Permisos y Reportaje – NOAA Fisheries Video Gallery. https://videos.fisheries.noaa.gov/detail/video/6347778430112/permiso-comercial-caribe%C3%B1o-de-botes-peque%C3%B1os-para-hms:-requisitos-de-permisos-y-reportaje?autoStart=true&q=shark

NOAA Fisheries (2023). Queen Conch. NOAA. https://www.fisheries.noaa.gov/species/queen-conch#:~:text=The%20queen%20conch%20occurs%20throughout,types%20of%20algae%20and%20epiphytes.

NOAA Fisheries (2022). “A Better Chance for Resilience”: Using Hungry Fish to Conserve Coral Reefs. NOAA. https://www.fisheries.noaa.gov/feature-story/better-chance-resilience-using-hungry-fish-conserve-coral-reefs#:~:text=Seaweed%2C%20or%20macroalgae%2C%20competes%20with,can%20overgrow%20and%20smother%20corals.

NOAA Fisheries (2022). Highly Migratory Species. NOAA. https://www.fisheries.noaa.gov/highly-migratory-species

NOAA Fisheries (2022). Impacts of Invasive Lionfish. NOAA. https://www.fisheries.noaa.gov/southeast/ecosystems/impacts-invasive-lionfish

NOAA Fisheries (2022). Understanding Atlantic Shark Fishing. NOAA. https://www.fisheries.noaa.gov/insight/understanding-atlantic-shark-fishing#:~:text=None%20of%20the%2043%20Atlantic,globally%E2%80%94are%20classified%20as%20threatened.

NOAA Fisheries (2021). Shark Identification Guide: Under the Highly Migratory Species (HMS) Commercial Caribbean Small Boat Permit. NOAA. https://media.fisheries.noaa.gov/2021-09/Caribbean-Shark-Placard-English-1-.pdf

NOAA Fisheries (2021). Understanding Essential Fish Habitat. NOAA. https://www.fisheries.noaa.gov/insight/understanding-essential-fish-habitat

NOAA Fisheries (2018). Laws & Policies: Endangered Species Act. NOAA. https://www.fisheries.noaa.gov/topic/laws-policies/endangered-species-act

NOAA Fisheries (2017). Shark Identification and Federal Regulations for the Recreational Fishery of the U.S. Atlantic, Gulf of Mexico and Caribbean. NOAA. https://www.fisheries.noaa.gov/resource/outreach-materials/shark-identification-placard

NOAA Office of Ocean Exploration and Research (2019). Exploring Puerto Rico’s Seamounts, Trenches, and Troughs.

NOAA Ocean Exploration. NOAA. https://oceanexplorer.noaa.gov/okeanos/explorations/ex1502/background/edu/purpose.html#:~:text=The%20Puerto%20Rico%20Trench%20is,water%20depths%20exceeding%208%2C400%20meters.

Nybakken, J. W. (2001). Marine Biology: An Ecological Approach. Bejamin Cummings. 

O

Ocean ART Project (2024). Sobrepesca. Universitat Politècnica de València, España. https://oceanartproject.blogs.upv.es/sobrepesca/

Ocean Exploration Trust (2017). Sharks of the Puerto Rico Trench – Nautilus Live [Video]. Nautilus Live. https://nautiluslive.org/video/2013/10/16/sharks-puerto-rico-trench

Office of Sustainable Fisheries Highly Migratory Species Management Division (2025). HMS Compliance Guide: Commercial Fishing – Guide for Complying with the Atlantic Tunas, Swordfish, Shark, and Billfish Regulations. 

Office of Sustainable Fisheries Atlantic Highly Migratory Species Management Division. (2023). HMS Compliance Guide: Recreational Fishing Guide for Complying with the Atlantic Tunas, Swordfish, Shark, and Billfish Regulations.

https://www.fisheries.noaa.gov/s3/2023-05/HMS-Recreational-Compliance-Guide-05-04-23.pdf

Officer, C. B., Ewing, J. I., Edwards, R. S., Johnson, H. R. (1957). Geophysical Investigations in the Eastern Caribbean: Venezuelan Basin, Antilles Island Arc, and Puerto Rico Trench. Bulletin of the Geological Society of America, 68, 359-378.

Oficina de Pesquerías Sostenibles: División de Manejo de Especies Altamente Migratorias (2025). Guía de Cumplimiento para HMS: Pesca Recreacional – Guía para el Complimiento de las Regulaciones para Atunes, Pez Espada, Pez de Pico, y Tiburón. https://www.fisheries.noaa.gov/s3/2025-08/Guia-de-Cumplimiento-Recreacional-de-HMS_Final_.pdf

Øgaard, B., Rölla, G., Ruben, J., Dijkman, T., Arends, J. (1988). Microradiographic study of demineralization of shark enamel in a human caries model. European Journal of Oral Sciences, 96(3), 209–211. https://doi.org/10.1111/j.1600-0722.1988.tb01545.x

Olán Martínez, C. (2014). Let’s Hunt Lionfish! NOAA Sea Grant. https://seagrant.noaa.gov/lets-hunt-lionfish/

O’Malley, M., Townsend, K. A., Hilton, P., Heinrichs, S., and Stewart, J. D. (2017). Characterization of the Trade in Manta and Devil Ray Gill Plates in China and South-east Asia through Trader Surveys. Aquatic Conservation: Marine and  Freshwater Ecosystems, 27, 394–413. https://doi.org/10.1002/aqc.2670

Orrell T, Informatics Office (2023). NMNH Extant Specimen Records (USNM, US). Version 1.67. National Museum of Natural History, Smithsonian Institution. Occurrence dataset. https://doi.org/10.15468/hnhrg3 recuperado de GBIF.org https://www.gbif.org/occurrence/1322822915

Osgood, G. J., Baum, J. K. (2015). Reef Sharks: Recent Advances in Ecological Understanding to Inform Conservation. Journal of Fishing Biology, 87, 1489-1523.

Ostrovsky, A. N., Lidgard, S., Gordon, D. P., Schwaha, T., Grigory Genikhovich, Ereskovsky, A. (2015). Matrotrophy and placentation in invertebrates: a new paradigm. Biological Reviews, 91(3), 673–711. https://doi.org/10.1111/brv.12189

Oxford University Press (2023). Oxford Languages. https://languages.oup.com/google-dictionary-en/

Oxford University Press (2023). Territorial Behavior. Oxford Reference. https://www.oxfordreference.com/display/10.1093/oi/authority.20110803103158702

P

Pacoureau, N., Carlson, J. K., Kindsvater, H. K., Rigby, C. L., Winker, H., Simpfendorfer, C. A., Charvet, P., Pollom, R. A., Barreto, R., C. Samantha Sherman, Talwar, B. S., Skerritt, D. J., U. Rashid Sumaila, Matsushiba, J. H., VanderWright, W. J., Yan, H. F., & Dulvy, N. K. (2023). Conservation successes and challenges for wide-ranging sharks and rays. Proceedings of the National Academy of Sciences of the United States of America, 120(5). https://doi.org/10.1073/pnas.2216891120

Padian, K. (2018, enero 18). Ostracoderm. Encyclopedia Britannica. https://www.britannica.com/animal/vertebrate/The-tetrapods

Pappas, K., Dunlap, K. (2011). Sensory Systems, Perception, and Learning | Shocking Comments: Electrocommunication in Teleost Fish. En A. P. Farrell (Ed.), Encyclopedia of Fish Physiology. Academic Press.

Park, H. B., Lam, Y. C., Gaffney, J. P., Weaver, J. C., Krivoshik, S. R., Hamchand, R., Pieribone, V., Gruber, D. F., Crawford, J. M. (2019). Bright Green Biofluorescence in Sharks Derives from Bromo-Kynurenine Metabolism. iScience, 19, 1277-1286. https://doi.org/10.1016/j.isci.2019.07.019

Parra, N. (2023, May 30). The Hidden Dangers of Shark Meat: Mislabeling, Sustainability, and Health Risks. Oceanic Preservation Society. https://www.opsociety.org/shark-meat#:~:text=The%20terms%20for%20shark%20meat,%2C%20rock%20salmon%2C%20and%20moki.

Parry, G. D. (1981). The Meanings of r- and K-Selection. Oecologia, 48, 260-264.

Parton, K. J., Godley, B. J., Santillo, D., Tausif, M., Omeyer, L. C. M., Galloway, T. S. (2020). Investigating the Presence of Microplastics in Demersal Sharks of the North-east Atlantic. Scientific Reports, 10, 12204. https://doi.org/10.1038/s41598-020-68680-1

Patrick, W. S., Spencer, P., Ormseth, O., Cope, J., Field, J., Kobayashi, D., Gedamke, T., Cortés, E., Bigelow, K., Overholtz, W., Link, J., Lawson, P. (2009). Use of Productivity and Susceptibility Indicesto Determine Stock Vulnerability, with Example Applications to Six U.S. Fisheries (NOAA Technical Memorandum NMFS-F/SPO-101). National Oceanic and Atmospheric Administration. 

Pawar, P. R., Shirgaonkar, S. S. & Patil, R. B. (2016). Plastic marine debris: Sources, distribution and impacts on coastal and ocean biodiversity. PENCIL Publication of Biological Sciences, 3(1): 40 – 54.

Pazmiño, D. A., Lynne van Herderden, Simpfendorfer, C. A., Junge, C., Donnellan, S. C., Hoyos-Padilla, M., Duffy, C., Huveneers, C., Gillanders, B. M., Butcher, P. A., Maes, G. E. (2019). Introgressive hybridisation between two widespread sharks in the east Pacific region. Molecular Phylogenetics and Evolution, 136, 119–127. https://doi.org/10.1016/j.ympev.2019.04.013

Pérez-Zayas, J. J., Mignucci-Giannoni, A. A., Toyos-González, G. M., Rosario-Delestre, R. J., Williams, Jr., E. H. (2002). Incidental predation by a largetooth cookiecutter shark on a Cuvier’s beaked whale in Puerto Rico. Aquatic Mammals, 28(2), 308-311.

Pillans, R. D., Stevens, J. D., Kyne, P. M., Salini, J. (2009). Observations on the Distribution, Biology, Short-Term Movements and Habitat Requirements of River Sharks Glyphis spp. in Northern Australia. Endangered Species Research, 10, 321-332. doi: 10.3354/esr00206

Pimiento, C., Ehret, D. J., MacFadden, B. J., Hubbell, G. (2010). Ancient Nursery Area for Extinct Giant Shark Megalodon from the Miocene of Panama. PLoS ONE, 5(5).

Pimiento, C., Leprieur, F., Silvestro, D., Lefcheck, J. S., Albouy, C., Rasher, D. B., Davis, M., Svenning, J.-C. ., Griffin, J. N. (2020). Functional diversity of marine megafauna in the Anthropocene. Science Advances, 6(16). https://doi.org/10.1126/sciadv.aay7650

Pirog A., Magalon H., Poirout T., Jaquemet S. (2019). Reproductive biology, multiple paternity and polyandry of the bull shark Carcharhinus leucas. Journal of Fish Biology, 95(5), 1195-1206. doi: 10.1111/jfb.14118.

Post, E., Thompson, J. N. (2017). Population Ecology. Encyclopædia Britannica. https://www.britannica.com/science/population-ecology

Pratt, H. (1979). Reproduction in the Blue shark, Prionace glauca. Fishery Bulletin, 77(2), 445-470.

Prevost, J. F. (1995). Nurse Sharks. ABDO.

Products Containing Sharks. (2013). Rob Stewart Sharkwater Foundation. https://www.robstewartsharkwaterfoundation.org/articles/products-containing-sharks

R

Ramos García, J. (2023). ¡Come pez león!: El recetario. Programa de Colegio Sea Grant de la Universidad de Puerto Rico.

Ramos García, J., Maldonado Rivera, A. (2013). No dejes que te coma el León. ¡Cómetelo tú! Programa de Colegio Sea Grant de la Universidad de Puerto Rico; Centro Interdisciplinario de Estudios del Litoral.

Real Academia Española (2024). Diccionario de la Lengua Española. https://dle.rae.es/

Rediker, M. (2008) History from Below the Water Line: Sharks and the Atlantic Slave Trade. Atlantic Studies, 5(2), 285-297, doi: 10.1080/14788810802149758

Reglamento de Pesca de Puerto Rico, Reglamento Núm. 7949-2010 (2010). https://www.drna.pr.gov/reglamentos/reglamento-num-7949/

Riaz, N. (2021). The Truth About Shark Culling & the Devastating Environmental Impacts. Conservation Made Simple. https://www.conservationmadesimple.org/blog/the-truth-about-shark-culling-the-devastating-environmental-impacts

Rivera-López, J. (1970). Studies on Biology of the Nurse Shark, Ginglymostoma cirratum Bonnaterre and the Tiger Shark, Galeocerdo cuvier Peron and Le Sueur. Universidad de Puerto Rico en Mayagüez.

Robertson, D. R., Van Tassell, J. (2023). Shorefishes of the Greater Caribbean. Smithsonian Tropical Research Institute. https://biogeodb.stri.si.edu/caribbean/en/pages

Robertson, D. R., Peña, E. A., Posada, J. M., Claro, R., Estape, A. C. (2023). Peces Costeros del Gran Caribe: sistema de Información en línea. Version 3.0 Instituto Smithsonian de Investigaciones Tropicales, Balboa, República de Panamá. https://biogeodb.stri.si.edu/caribbean/es/ 

Robins R (2023). UF FLMNH Ichthyology. Version 117.394. Florida Museum of Natural History. Occurrence dataset. https://doi.org/10.15468/8mjsel recuperado de GBIF.org https://www.gbif.org/occurrence/624129343

Robins R (2023). UF FLMNH Ichthyology. Version 117.394. Florida Museum of Natural History. Occurrence dataset. https://doi.org/10.15468/8mjsel recuperado de GBIF.org https://www.gbif.org/occurrence/2989399302

Robins R (2023). UF FLMNH Ichthyology. Version 117.394. Florida Museum of Natural History. Occurrence dataset. https://doi.org/10.15468/8mjsel recuperado de GBIF.org https://www.gbif.org/occurrence/624129200

Rod and Reel. (2013). Law Insider. https://www.lawinsider.com/dictionary/rod-and-reel#:~:text=Rod%20and%20reel%20means%20a,is%20wound%20around%20a%20barrel.

Rodríguez-Ferrer, G., Wetherbee, B. M., Schärer, M., Lilyestrom, C., Zegarra, J. P., Shivji, M. (2017). First Record of the Megamouth Shark, Megachasma pelagios, (family Megachasmidae) in the Tropical Western North Atlantic Ocean. Marine Biodiversity Records, 10(20).

Roff, G., Brown, C. J., Priest, M. A., Mumby, P. J. (2018). Decline of coastal apex shark populations over the past half century. Communications Biology, 1(223).

Roff, G., Doropoulos, C., Rogers, A., Bozec, Y.-M., Krueck, N. C., Aurellado, E., Priest, M., Birrel, C., Mumby, P. J. (2016). The Ecological Role of Sharks on Coral Reefs. Trends in Ecology & Evolution, 31(5), 395–407. doi:10.1016/j.tree.2016.02.01

Rull, V. (2020). Humans: occupation and humanization of the planet. Quaternary Ecology, Evolution, and Biogeography, 173–221. https://doi.org/10.1016/b978-0-12-820473-3.00005-1

Russell G. M. , Gutherz E. J.,  Barans C. A. (1988). Evaluation of demersal longline gear off South Carolina and Puerto Rico with emphasis on deep-water reef fish stocks. Marine Fisheries Review, 50, 26–31.

Rutledge, K., McDaniel, M., Teng, S., Hall, H., Ramroop, T., Sprout, E., Hunt, J., Boudreau, D., Costa, H. (2024). Food Chain. National Geographic: Education. https://education.nationalgeographic.org/resource/food-chain/

Ryan, L. A., Slip, D. J., Chapuis, L., Collin, S. P., Gennari, E., Hemmi, J. M., How, M. J., Huveneers, C., Peddemors, V. M., Tosetto, L., Hart, N. S. (2021). A Shark’s Eye View: Testing the ‘Mistaken Identity Theory’ Behind Shark Bites on Humans. The Royal Society, 18(183). https://doi.org/10.1098/rsif.2021.0533

S

Sanders, R. (2022, septiembre 15). How Saturn got its rings. University of California. https://www.universityofcalifornia.edu/news/how-saturn-got-its-rings#:~:text=Militzer%2C%20UC%20Berkeley%20professor%20of,too%20close%20to%20the%20planet.

Sapag, C. (2021). ¿Qué es la pesca de arrastre de fondo? Oceana Chile. https://chile.oceana.org/blog/que-es-la-pesca-de-arrastre-de-fondo/#:~:text=marino%20en%20s%C3%AD.-,Las%20redes%20de%20arrastre%20de%20media%2Dagua%2C%20o%20pel%C3%A1gicas%2C,el%20h%C3%A1bitat%20donde%20%C3%A9stas%20operan.

Saville, K., Lindley, A., Maries, E., Carrier, J., Pratt, H. (2002). Multiple Paternity in the Nurse Shark, Ginglymostoma cirratum. Environmental Biology of Fishes, 63, 347-351. doi:10.1023/A:1014369011709

Scaradozzi, D. Palmieri, G. Costa, D. Pinelli, A. (2017). BCF swimming locomotion for autonomous underwater robots: a review and a novel solution to improve control and efficiency. Ocean Engineering, 130, 437–453. doi:10.1016/j.oceaneng.2016.11.055

Schärer-Umpierre, M., Franqui-Rivera, G. (2022). First confirmed location of Carcharhinus signatus in Puerto Rico. Journal of the Marine Biological Association of the United Kingdom, 1-4. https://doi.org/10.1017/ S0025315422000881

Schärer-Umpierre, M., Ruiz, H., Zayas-Santiago, C., Bond, M. E. (2020). Coral reef elasmobranchs in Puerto Rico’s fishery and MPA effects. Proceedings of the 73rd Gulf and Caribbean Fisheries Institute, 51-53. 

Scheid P., Pelster B., Kobayashi H. (1990). Gas exchange in the fish swimbladder. Advances in Experimental Medicine and Biology, 277, 735-42. doi: 10.1007/978-1-4684-8181-5_8

Schneider, B. (2013, Jun 13). An Interconnected Environment and Economy: Shark Tourism in Palau. Scientific Research Diving at USC Dornsife. https://uscdiving.wordpress.com/2013/06/13/originally-published-at-scientificamerican-comby-brenna-schneideras-a-small/

Sea Grant Puerto Rico (2015). Vedas en aguas de Puerto Rico. https://seagrantpr.org/wp-content/uploads/2015/07/tabla_de_vedas.pdf

Secretaría de Agricultura y Desarrollo Rural (2026). ¿Por qué se establecen las vedas? Gobierno de México. https://www.gob.mx/agricultura/articulos/por-que-se-establecen-las-vedas?idiom=es#:~:text=Las%20vedas%20que%20son%20los,permitir%20su%20reproducci%C3%B3n%20y%20subsistencia.

Shark Senses. (2023). Smithsonian Ocean. https://ocean.si.edu/ocean-life/sharks-rays/shark-senses

Shark Steak Recipe: Common Blue Shark Steak. (2021). Glutenfreemama. https://glutenfreemama.ru/en/steik-akuly-kak-prigotovit-recept-steik-iz-obychnoi-goluboi-akuly/

Shiffman, D. (2022). Why sharks matter? Johns Hopkins University Press.

Simpfendorfer, C. A., Heupel, M. R., White, W. T., Dulvy, N. K. (2011). The Importance of Research and Public Opinion to Conservation Management of Sharks and Rays: A Synthesis. Marine and Freshwater Research, 62, 518-527.

Skomal, G. B., Zeeman, S. I., Chisholm, J. H., Summers, E. L., Walsh, H. J., McMahon, K. W., Thorrold, S. R. (2009). Transequatorial Migrations by Basking Sharks in the Western Atlantic Ocean. Current Biology, 19(12), 1019-1022.

Smith, L. (2023, diciembre 12). Pregnant megamouth shark seen for 1st time after female washes up dead with 7 pups. Live Science. https://www.livescience.com/animals/sharks/pregnant-megamouth-shark-seen-for-1st-time-after-female-washes-up-dead-with-7-pups

Smith, W. L., Stern, J. H., Girard, M. G., Davis, M. P. (2016). Evolution of Venomous Cartilaginous and Ray-Finned Fishes. Integrative and Comparative Biology, 56(5), 950–961. https://doi.org/10.1093/icb/icw070

Soler-Figueroa, B. M., Otero, E. (2014). The Influence of Rain Regimes and Nutrient Loading on the Abundance of Two Dinoflagellate Species in a Tropical Bioluminescent Bay, Bahía Fosforescente, La Parguera, Puerto Rico. Estuaries and Coasts, 38(1), 84–92. https://doi.org/10.1007/s12237-014-9827-0

Soto, S., Pintó, J. (2010). Delineation of natural landscape units for Puerto Rico. Applied Geography, 30(4), 720-730. https://doi.org/10.1016/j.apgeog.2010.01.010

Speedo USA (2026). What Is A Rash Guard? Speedo International Limited, Speedo.com. https://us.speedo.com/faq/what-is-a-rash-guard.list

Spiers, E. K. A., Stafford, R., Ramirez, M., Vera Izurieta, D. F., Cornejo, M., Chavarria, J. (2016). Potential Role of Predators on Carbon Dynamics of Marine Ecosystems as Assessed by a Bayesian Belief Network. Ecological Informatics, 36, 77-83. https://doi.org/10.1016/j.ecoinf.2016.10.003

Stanley, M. (2023). Marine Ecosystems. National Geographic. https://education.nationalgeographic.org/resource/marine-ecosystems/

Statistical Software Package: An overview (2023). ScienceDirect Topics. https://www.sciencedirect.com/topics/computer-science/statistical-software-package

Stevens, J. D., Bonfil, R., Dulvy, N. K., Walker, P. A. (2000). The Effects of Fishing on Sharks, Rays and Chimeras (Chondrichthyans), and the Implications for Marine Ecosystems. ICES Journal of Marine Science, 57, 476 – 494.

Straube, N., Lampert, K. P., Geiger, M. F., Weiß, J. D., Kirchhauser, J. X. (2016). First record of second-generation facultative parthenogenesis in a vertebrate species, the whitespotted bambooshark Chiloscyllium plagiosum. Journal of Fish Biology, 88(2), 668–675. doi:10.1111/jfb.12862

Subsecretaría de Pesca y Acuicultura (2018). Especialista de Subpesca explica qué son las vedas y cuál es su propósito. SUBPESCA: Difusión. https://www.subpesca.cl/portal/difusion/Noticias/100030:Especialista-de-Subpesca-explica-que-son-las-vedas-y-cual-es-su-proposito#:~:text=Las%20vedas%20son%20medidas%20de,departamento%20de%20pesquer%C3%ADas%2C%20Javier%20Rivera.

Sullivan, J. (2013). Top 10 Whale Shark Facts. Oceana USA. https://usa.oceana.org/blog/top-10-whale-shark-facts/#:~:text=Whale%20sharks%20are%20extremely%20unusual,size%20of%20a%20manhole%20cover!

Sustainable Fisheries UW (2019, abril 28). Sustainable Seafood 101: Fishery Management. University of Washington scientists; Sustainable Fisheries UW. https://sustainablefisheries-uw.org/seafood-101/fishery-management/#:~:text=Fishing%20management%20is%20the%20process%20of%20regulating,fisheries%20are%20regulated%20by%20the%20Magnuson%2DStevens%20Act.

Suuronen, P., He, P., Pol, M., Graham, N., Reid, D. (2014). Selective Fishing, Non-Selective Fishing and Balanced Fishing: Concept, Consequences and Challenges. American Fisheries Society 144th Annual Meeting. 

T

TED-ed (2023). Why did Megalodon go extinct? – Jack Cooper and Catalina Pimiento. [Video]. YouTube. https://www.youtube.com/watch?v=6LGCk08zMbg  

The Editors of Encyclopaedia Britannica (2023). Encylopaedia Britannica. https://www.britannica.com

The Importance of Sharks. (2023). Oceana Europe. https://europe.oceana.org/importance-sharks-0/

The Smithsonian Institution’s Human Origins Program (2024, julio 9). Introduction to Human Evolution. Smithsonian National Museum of Natural History. https://humanorigins.si.edu/education/introduction-human-evolution#:~:text=Full%20Image,evolution%20occurred%20on%20that%20continent.‌

Tirnadmin (2016). Cocos Island Cleaning Stations: Providing Sharks with a Healthy Spa Resort. Turtle Island Restoration Network. https://seaturtles.org/cleaning-stations/#:~:text=Some%20of%20the%20most%20fascinating,species%2C%20often%20by%20smaller%20fish.

Tribe (2021). DBpedia. https://dbpedia.org/page/Tribe_(biology)

Tilley, A., López-Angarita, J., Turner, J. R. (2013) Diet Reconstruction and Resource Partitioning of a Caribbean Marine Mesopredator Using Stable Isotope Bayesian Modelling. PLoS ONE 8(11): e79560. https://doi.org/10.1371/journal.pone.0079560

Tytell, E. D., Iman Borazjani, Sotiropoulos, F., T. Vernon Baker, Anderson, E. J.,  Lauder, G. V. (2010). Disentangling the Functional Roles of Morphology and Motion in the Swimming of Fish. Integrative and Comparative Biology, 50(6), 1140–1154. https://doi.org/10.1093/icb/icq057

U

Unidad de la Marejada Ciclónica (s.f.). Introducción a la Marejada Ciclónica. Centro Nacional de Huracanes; NOAA.

Unión Internacional para la Conservación de la Naturaleza (2025). The IUCN Red List of Threatened Species. Version 2025-2. https://www.iucnredlist.org. 

United States Environmental Protection Agency (2025, julio 2). Microplastics Research. US EPA. https://www.epa.gov/water-research/microplastics-research#:~:text=Secondary%20microplastics%20are%20plastic%20particles,surface%20water%20and%20aquatic%20species.

U.S. Food and Drug Administration (2020). FDA/EPA 2004 Advice on What You Need to Know About Fish. https://www.fda.gov/food/environmental-contaminants-food/fdaepa-2004-advice-what-you-need-know-about-mercury-fish-and-shellfish

V

Valdés Pizzini, M., Schärer Umpierre, M. T. (2011). Una mirada al mundo de los pescadores en Puerto Rico. Programa de Colegio Sea Grant de la Universidad de Puerto Rico; Centro Interdisciplinario de Estudios del Litoral.

Verbeke, R. (2025). A 400-million-year-old placoderm reveals how the first dentitions were structured. Institute of Natural Sciences. https://www.naturalsciences.be/en/science/news/a-400-million-year-old-placoderm-reveals-how-the-first-dentitions-were-structured

Vianna, G. M. S., Meekan, M. G., Pannell, D., Marsh, S., Meeuwig, J. J. (2010). Wanted Dead or Alive?: The relative value of reef sharks as a fishery and an ecotourism asset in Palau. Australian Institute of Marine Science; University of Western Australia. https://www.issuelab.org/resources/26562/26562.pdf

Villagómez Vélez, S. I. (2016). Comportamiento alimentario de Tiburón ballena (Rhincodon typus) ante la presencia de turistas en el norte Caribe mexicano. Universidad Nacional Autónoma de México. https://ru.dgb.unam.mx/bitstream/20.500.14330/TES01000741454/3/0741454.pdf

W

Waggoner, B. (2000, julio 22). Introduction to the Placodermi. University of California Museum of Paleontology. https://ucmp.berkeley.edu/vertebrates/basalfish/placodermi.html‌

Walker, T. I. (2005). General biology of Chondrichthyan fishes. Marine Education Society of Australasia. http://www.mesa.edu.au/Seaweek2005/pdf_senior/is01.pdf

Watson and the Shark. (2013). The National Gallery of Arts. https://www.nga.gov/collection/art-object-page.46471.html

Weigmann, S., Gon, O., Leeney, R. H., Barrowclift, E., Berggren, P., Jiddawi, N., Temple, A. J. (2020). Revision of the sixgill sawsharks, genus Pliotrema (Chondrichthyes, Pristiophoriformes), with descriptions of two new species and a redescription of P. warreni Regan. PLOS ONE, 15(3), e0228791. https://doi.org/10.1371/journal.pone.0228791

Welten M., Smith M. M., Underwood C., Johanson Z. 2015 Evolutionaryorigins and development of saw-teeth on the sawfish and sawshark rostrum (Elasmobranchii; Chondrichthyes).Roya Society Open Science, 2: 150189. http://dx.doi.org/10.1098/rsos.1501

Wetherbee, B. M., Crow, G. L., Lowe, C. G. (1996). Biology of the Galapagos shark, Carcharhinus galapagensis, in Hawai’i. Environmental Biology of Fishes, 45, 299–310. https://doi.org/10.1007/BF00003099

Wetzel, C. (2020). How some animals have ‘virgin births’: Parthenogenesis explained. National Geographic. https://www.nationalgeographic.com/animals/article/parthenogenesis-how-animals-have-virgin-births

What Animals Eat Sharks? (2023). American Oceans. https://www.americanoceans.org/facts/what-animals-eat-sharks/#:~:text=in%20many%20areas.-,Osprey,eat%20small%20sharks%20like%20dogfish.

White, J. A., Burgess, G. H., Nakatsukasa, M., Hudson, M. J., Pouncett, J., Kusaka, S., Yoneda, M., Yamada, Y., Schulting, R. J. (2021). 3000-year-old shark attack victim from Tsukumo shell-mound, Okayama, Japan. Journal of Archaeological Science: Reports, 38, 103065. https://doi.org/10.1016/j.jasrep.2021.103065

Williams, J. J., Papastamatiou, Y. P., Caselle, J. E., Bradley, D., Jacoby, D. M. P. (2018). Mobile Marine Predators: an Understudied Source of Nutrients to Coral Reefs in an Unfished Atoll. Proceedings of the Royal Society B, 285, 20172456. http://dx.doi.org/10.1098/rspb.2017.2456

Wilson, D. E., Jones, J. K., Armstrong, D. M. (2019). Mammal: Animal. In Encyclopædia Britannica. https://www.britannica.com/animal/mammal

Wilson, S. G. (2004). Basking sharks (Cetorhinus maximus) Schooling in the Southern Gulf of Maine. Fisheries Oceanography, 13(4), 283–286. doi:10.1111/j.1365-2419.2004.00292.x

Wong, K. (2009, Dec 28). Full Body Swimsuit Now Banned for Professional Swimmers. ABC News Network. https://abcnews.go.com/Politics/full-body-swimsuit-now-banned-professional-swimmers/story?id=9437780

World Register of Introduced Marine Species (2021). Open Ocean Habitat. https://www.coastalwiki.org/introduced/Open_ocean_habitat#:~:text=The%20open%20oceans%20or%20pelagic,surface%20to%20the%20abyssal%20depths

World Wide Fund For Nature (2020). Five reasons why oceanic sharks and rays matter to the world and people! https://sharks.panda.org/news-blogs-updates/latest-news/five-reasons-why-oceanic-sharks-and-rays-matter-to-the-world-and-people

WoRMS Editorial Board (2025). World Register of Marine Species. https://www.marinespecies.org 

WWF (2023). Tuna. World Wildlife Fund. https://www.worldwildlife.org/species/tuna#:~:text=Their%20specialized%20body%20shape%2C%20fins,incredible%20distances%20as%20they%20migrate.‌

Y

You. J. (2014). Mercury levels in surface ocean have tripled. Science. https://www.science.org/content/article/mercury-levels-surface-ocean-have-tripled#:~:text=In%20gold%20mining%2C%20miners%20use,rain%20deposits%20it%20into%20oceans.

Young, C. N., Carlson, J., Hutchinson, M., Kobayashi, D., McCandless, C., Miller, M. H., Teo, S., T. Warren. (2016). Status Review Report: Common Thresher Shark (Alopias vulpinus) and Bigeye Thresher Shark (Alopias superciliosus). Final Report to National Marine Fisheries Service; Office of Protected Resources.

Z

Zarza, L. F. (2019). ¿Qué son las aguas grises? iAgua. https://www.iagua.es/respuestas/que-son-aguas-grises

#

50 CFR Part 635 – Atlantic Highly Migratory Species. (2024). Ecfr.gov. https://www.ecfr.gov/current/title-50/chapter-VI/part-635